After Rocky's comments, I was prompted to call Alain Desjardins, the gentleman I bought the SR block from, to ask him if the caps were on the block when he got it, and if not, where they came from. I got a really helpful detailed story:
He did not know the man he bought the block from. The seller had a few Pontiac blocks for sale. Alain asked a lot of questions without getting any very helpful answers about the block, it's condition, casting numbers etc. but he decided to go and see it anyway. When he arrived he found three Pontiac blocks stored in a small, old, unheated warehouse, along with a lot of junk. When he first got there, none of the blocks had the crank or the main caps installed.
The three blocks were a 400, the SR 455, and a YY 455 with a casting date of F233 - July 23, 1973. .
After looking through the pile of junk, they located two crankshafts, one with 3.00 journals and one with 3.25 journals - so for a 400 and for a 455. Both of these cranks were rusty, dirty and scratched and were not of much interest. The seller was unable to give Alain any more information. Looking through the junk, they found a small box with the main caps in it. Alain thought that of the blocks that were there, they had to be from the SR block, since they couldn't be for the 400 because of the difference in the journal size, and the other 455 block was a '73, and the only 4 bolt main block in 1973 was the 455 SD, and the '73 block was obviously not a Super Duty. So he concluded that the main caps probably went with the SR block, but they weren't on it when he got there, and the seller didn't remember if they were from that block or not or where they came from. At the time, Alain did not know that all SR blocks dated 1972 and later came with 4 bolt mains; presumably, if he had known that, it would have reinforced his conclusion.
When he advertised the SR block for sale, he made no claim that main caps were original to the block, just that they were available to be bought with the block. He even suggested that they were not, and that the block should be line bored. He told me that since he couldn't prove for sure that the caps came with the block, he didn't want to risk misrepresenting it. Based on these facts, though (combined with Rocky's comments above) I am virtually certain that the main caps ARE original to the block, and I am absolutely certain that Alain Desjardins (of Quebec City, Quebec, if you happen to be dealing with him - he often sells Pontiac parts) is a very honest man.
And of course, Rocky nailed it from the beginning. I am really glad that he asked why I would think that the caps weren't original to the block, and that I got the whole story from Alain!